type specimen holotype Paleogene Oligocene North European Plain FEU-1027498
|  | identified by M. Hesemann Paleogene Oligocene Chattian North European Plain Kobrow FEU-1001051
|  | identified by M. Hesemann Paleogene Oligocene Chattian North European Plain Kobrow FEU-1001052
|  | identified by M. Hesemann Paleogene Oligocene Chattian North European Plain Kobrow FEU-1001053
|  | identified by M. Hesemann Paleogene Oligocene Chattian North European Plain Ahnetal FEU-1027692
|  |