type specimen lectotype Quaternary Holocene recent central North Pacific Challenger Station 241 FEU-1003825
| | identified by scientist Quaternary Holocene recent central North Pacific Challenger Station 241 FEU-1003188
| | identified by scientist Quaternary Holocene recent off Argentina Challenger Station 323 FEU-1003189
| | identified by scientist Quaternary Holocene recent central North Pacific Challenger Station 246 FEU-1003190
| | identified by scientist Quaternary Holocene recent unknown FEU-1011060
| |