type specimen holotype Paleogene Oligocene unknown FEU-1022379
| | identified by scientist Neogene Miocene Burdigalian Pannonian Basin Matraverebely FEU-1002826
| | identified by M. Hesemann Paleogene Oligocene Chattian North European Plain Kassel FEU-1009964
| | identified by M. Hesemann Paleogene Oligocene Chattian North European Plain Kassel FEU-1009965
| | identified by M. Hesemann Paleogene Oligocene Chattian North European Plain Kassel FEU-1009966
| |